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Instrumental records indicate a century-long trend towards drying over western North America and wetting over eastern North America. A continuation of these trends into the future would have significant hydroclimatic and socioeconomic consequences in both the semi-arid Southwest and humid East. Using tree-ring reconstructions and hydrologic simulations of summer soil moisture, we evaluate and contextualize the modern summer aridity gradient within its natural range of variability established over the past 600 years and evaluate the effects of observed and anthropogenic precipitation, temperature, and humidity trends. The 2001–2020 positive (wet east-dry west) aridity gradient was larger than any 20 year period since 1400 CE, preceded by the most negative (wet west-dry east) aridity gradient during 1976–1995, leading to a strong multi-decade reversal in aridity gradient anomalies that was rivaled only by a similar event in the late-16th century. The 2001–2020 aridity gradient was dominated by long-term summer precipitation increases in the Midwest and Northeast, with smaller contributions from more warming in the West than the East and spring precipitation decreases in the Southwest. Multi-model mean climate simulations from Coupled Model Intercomparison Project 6 experiments suggest anthropogenic climate trends should not have strongly affected the aridity gradient thus far. However, there is high uncertainty due to inter-model disagreement on anthropogenic precipitation trends. The recent strengthening of the observed aridity gradient, its increasing dependence on precipitation variability, and disagreement in modeled anthropogenic precipitation trends reveal significant uncertainties in how water resource availability will change across North America in the coming decades.

Rapid drought intensification, or flash droughts, is often driven by anomalous atmospheric ridging and can cause severe and complex impacts on water availability and agriculture, but the full range of variability of such events in terms of intensity and frequency is unknown. New tree‐ring reconstructions of May–July mid‐tropospheric ridging and soil moisture anomalies back to 1500 CE in the central United States—a hotspot for flash drought—suggest that over the last five centuries, anomalies in these two variables combined to indicate flash‐drought conditions in ∼17% of years and exceptionally severe flash drought in ∼4% of years, similar to frequencies in recent decades. However, over one‐third of all inferred exceptional flash droughts occurred since 1900, suggesting the 20th century was highly flash‐drought prone. These results may guide future work to diagnose the roles of external, oceanic, and land‐surface forcing of warm‐season atmospheric circulation and hydroclimate over North America.

Primary production is the entry point of energy and carbon into ecosystems, but modeling responses of primary production to “environmental stress” (i.e., reductions of primary production from nonoptimal environmental conditions) remains a key challenge and source of uncertainty in our understanding of Earth's carbon cycle. Here we develop an approach for estimating annual “environmental stress” from tree rings based on the proportion of the optimal diameter growth rate (from species‐specific allometric equations) that is realized in a given year. We assessed climatic, topographic, and soil drivers of environmental stress, as well as their interactions, using both empirical model experiments and linear mixed effect models. Climate gradients and interannual climate variability dominated spatial and temporal variability of environmental stress in much of the western United States, where the tree‐ring environmental stress index was positively correlated with antecedent climatic water balance (precipitation minus potential evapotranspiration) and negatively correlated with temperature and vapor pressure deficit. Excluding topographic and soil information from empirical models reduced their ability to capture spatial gradients in environmental stress, particularly in the eastern United States, where growth was not as strongly limited by climate. Mean climate conditions and topographic characteristics had significant interaction effects with the climatic water balance, indicating an increasing importance of winter moisture for warmer and drier sites and as elevation and topographic wetness index increased. These results suggest that including effects of antecedent climate (particularly in dry regions) and site topographic and soil characteristics could improve parameterization of environmental stress effects in primary production models.

Much of the eastern United States experienced increased precipitation over the twentieth century. Characterizing these trends and their causes is critical for assessing future hydroclimate risks. Here, U.S. precipitation trends are analyzed for 1895–2016, revealing that fall precipitation in the southeastern region north of the Gulf of Mexico (SE-Gulf) increased by nearly 40%, primarily increasing after the mid-1900s. Because fall is the climatological dry season in the SE-Gulf and precipitation in other seasons changed insignificantly, the seasonal precipitation cycle diminished substantially. The increase in SE-Gulf fall precipitation was caused by increased southerly moisture transport from the Gulf of Mexico, which was almost entirely driven by stronger winds associated with enhanced anticyclonic circulation west of the North Atlantic subtropical high (NASH) and not by increases in specific humidity. Atmospheric models forced by observed SSTs and fully coupled models forced by historical anthropogenic forcing do not robustly simulate twentieth-century fall wetting in the SE-Gulf. SST-forced atmospheric models do simulate an intensified anticyclonic low-level circulation around the NASH, but the modeled intensification occurred farther west than observed. CMIP5 analyses suggest an increased likelihood of positive SE-Gulf fall precipitation trends given historical and future GHG forcing. Nevertheless, individual model simulations (both SST forced and fully coupled) only very rarely produce the observed magnitude of the SE-Gulf fall precipitation trend. Further research into model representation of the western ridge of the fall NASH is needed, which will help us to better predict whether twentieth-century increases in SE-Gulf fall precipitation will persist into the future.

Projected changes in temperature and drought regime are likely to reduce carbon (C) storage in forests, thereby amplifying rates of climate change. While such reductions are often presumed to be greatest in semi‐arid forests that experience widespread tree mortality, the consequences of drought may also be important in temperate mesic forests of Eastern North America (ENA) if tree growth is significantly curtailed by drought. Investigations of the environmental conditions that determine drought sensitivity are critically needed to accurately predict ecosystem feedbacks to climate change. We matched site factors with the growth responses to drought of 10,753 trees across mesic forests ofENA, representing 24 species and 346 stands, to determine the broad‐scale drivers of drought sensitivity for the dominant trees inENA. Here we show that two factors—the timing of drought, and the atmospheric demand for water (i.e., local potential evapotranspiration;PET)—are stronger drivers of drought sensitivity than soil and stand characteristics. Drought‐induced reductions in tree growth were greatest when the droughts occurred during early‐season peaks in radial growth, especially for trees growing in the warmest, driest regions (i.e., highestPET). Further, mean species trait values (rooting depth and ψ₅₀) were poor predictors of drought sensitivity, as intraspecific variation in sensitivity was equal to or greater than interspecific variation in 17 of 24 species. From a general circulation model ensemble, we find that future increases in early‐seasonPETmay exacerbate these effects, and potentially offset gains in C uptake and storage inENAowing to other global change factors.